Betas- Beta's are thought of as highly wolves, they help lead hunts with the alphas and watch fights to find the winner. They can't promote but wolves try to please them. They are of second-in-command and are strong too. Sometimes in really rare cases a male beta can be somewhat larger or strong than a femme alpha. But no femme beta is larger or stronger than an alpha.
Sigmas- Sigmas are still a permanent ranking wolves but they don't do much. In the hunt they run with the pack but have the power to push them in line if they fail. Sigma's also get a good meal after a hunt. Sigmas are ussually of in-pack relations as betas and alphas. Usually a wolf that joined from the outside won't go as high as a Sigma, unless they show wonderful tactics and attitude. Reason being: Wolves with In-Pack relation have more to lean back on, if you are a wolf of alpha blood you are considered somewhat like your parents, then given some leeway to raise yourself a rank. But some woulds squander the family heritage thus ruining it for themselves.
Selsa- Selsa's pretty much hold the same role as Sigmas, they just don't get much respect for it. A selsa will watch the pack though. Selsas are somewhat strong, but ussually loyalty gets them here. But in most cases a wolf must be a good fighter to be a Selsa, incase anything bad happens they are needed to fight.
Delta & Gammas- They hold the same role, they play more of an 'older sibling' type of a stance. Ussually when a large group of pups grows up the dominant one assumes this rank and the others spread out.
Epsilons- These are out of the permanent ranking zone and are into the pack wolves. They don't do much, in some cases dominant pups assume this role emmediatly after growing up. They are strong and are respected, but not considered high. They have no jobs, they just are the noble dudley-do-rights.
Zetas- They are the first assumed rank most wolves will get. Since iotas are ussually gotten by anyone. They are the rank that just did something good and are finding their feet in that rank. In some cases zetas may be deranked epsilons. Whom the alphas thought them merely not ready to assume responsibility.
Iota- Iotas are usually the bulk of the pack. They are most of the pups that just became adults, newer members, and disrespectful wolves that weren't as bad to be omegas. They are slightly like omegas, but given more leeway.
Omegas- these are the disrespectful wolves. They have to respect everyone else in the pack. There is usually 1 or 2 omegas in a pack and they are given no mercy, hence the nickname 'scapegoat'. In a pack the wolves may practice their rank and power on the omegas, but wolves if they tried can easily get out of this rank. Unless they were horribly bad.
now I hope this helps you understand the pack life and how you get certain ranks. All in all no rank has a certain job, they just have a certain level of respect.beware,lower ranks,disrespect higher ranks.....ouch.
another tip;once a wolf goes up to a high position,like beta,and they lose a fight,they either are put at a very low rank,or kicked out of the pack.