The main purpose of the
digestive system is to take in food, digest it, and absorb nutrients into the body.
Digestion in the horse takes place through muscular action, enzyme action, and bacterial
fermentation. The digestive system is composed of the alimentary canal and the accessory
organs, which include the teeth, salivary glands, liver, and pancreas. The alimentary
canal is approximately 100 feet long, and starts at the mouth, continues to the pharynx,
esophagus, stomach, small intestine, large intestine, and ends at the anus.
The Mouth and Pharynx
The process of digestion begins at the mouth with the lips and incisors grasping food. The
molars then grind the food while mixing it with saliva. This alkaline saliva is produced
from the paratoid, submaxmillary and sublingual glands. The average mature horse produces
about 10 gallons of saliva per day. This saliva acts as a digestive juice on the sugars
and starches in the feed and lubricates it for swallowing. The tongue flips balls of food
back to the pharynx once it has thoroughly chewed. The pharynx is a muscular passage to
the esophagus. TOP
The Esophagus
The esophagus is a 5-foot long tube with circular muscles, which force the food and water
down to the stomach. These muscular waves of constriction are known as peristalsis. The
angle at which the esophagus meets the stomach is such that excessive gastric buildup
pushes the membrane flaps of the opening closed. This makes vomiting in the horse
extremely difficult. A horse that is vomiting may have ruptured is stomach lining. TOP
The Stomach
The stomach is J-shaped organ with the openings to the esophagus and small intestine close
together. Digestion is best achieved when the stomach is only 2/3 full, thus the
importance of several smaller meals, as opposed to one or two large feeding. Cells in the
lining primarily secrete hydrochloric acid and the enzyme pepsinogen. The hydrochloric
acid acts to change pepsinogen to pepsin, a protein digester, and it dissolves mineral
matter. Food enters the stomach in layers and stays until the 2/3 full. At this point, is
pushed out to the small intestine. During a large meal, some of the food being pushed out
will not have had the chance to be totally digested. TOP
The Small Intestine
The small intestine is about 70 feet long and 2 to 3 inches in diameter. It is divided
into three parts for identification, the duodenum, the jejunum, and the ileum. The inner
lining is comprised of specialized mucosal cells arranged into villi, which vastly
increases the absorptive surface area. These cells perform the major part of the enzymatic
breakdown of feed. Proteins, sugars, fats are absorbed through the villi and enter the
blood stream.
The three main types of enzymes in the small intestine are:
protein-digesting, carbohydrate-digesting, and fat-digesting. The protein-digesting
enzymes, trypsin and chrymrotrypsin, are created in a non-active form in the pancreas and
become active protein-digesters in the small intestine. Trypsin and chymrotrypsin are not
totally effective unless preceded by the pepsin enzyme in the stomach. These enzymes
produce the final breakdown of proteins to amino acids. The carbohydrate-digesting enzyme
amylase is formed in the pancreas and acts on starches, changing them to maltose, a sugar.
This maltose and other 2 part sugars (disaccarides) are hydrolyzed, or broken down by
inserting water molecules, single part sugars (monosaccarides) and absorbed. Fat-digesting
enzymes are also formed in the pancreas, as well as the small intestine. The affect lipids
(fats) and break them down into fatty acids and monoglycerides by hydrolysis.
Muscular contractions mix the food in four different variations in
the small intestine. The first type of contraction is the contraction and relaxation of
various short segments. The second type is pendular: the swaying back and forth of the
small intestine. The third type of muscular contraction occurs in short propulsive
movements; weak waves of contraction, which move about 2 inches before stopping. The
contractions then pause for a moment before starting up where they left off. The final
type of contraction is paristalsis, which is not normal in the small intestine and results
in diarrhea-like symptoms. TOP
The Large Intestine
The large intestine is divided into four parts; the cecum, the large colon, the small
colon, and the rectum. The cecum is 4 feet long and contains mostly liquid. The food and
water ingested by the horse are well mixed here. The bacterial population breaks down the
food, resulting in the formation of fatty acids and some vitamins. The large colon is 10
to 12 feet long and 8 to 18 inches in diameter. Some digestion, mostly bacterial
fermentation and absorption occur here. The small colon is also about 10 to 12 feet in
length, but is only 3 to 4 inches in diameter. Most of the moisture is absorbed in the
colons. The remaining food matter is formed into balls at the end of the small colon. The
last foot of the large intestine is the rectum, where the waste is stored until its is
passed. TOP
The Liver and Pancreas
The liver is the largest gland in the body. It produces bile and processes the digested
proteins, sugars, minerals, and other food particles. It then regulates the particles for
immediate use or storage. The pancreas produces enzymes for the small intestine and
insulin, which is need for the metabolism of sugar in the blood stream. TOP